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For bird species in which females are the sole incubators, a male can increase the female's percentage of time spent on the nest (nest attentiveness) by feeding the incubating female on the nest (incubation feeding). Given the benefits of nest attentiveness and the potential ability of incubation feeding to increase attentiveness, the question arises of why species vary in their rate of incubation feeding. One hypothesis suggests that greater incubation feeding is required in harsher microclimates because there the cost of leaving eggs unattended is greater. Under this hypothesis, nest attentiveness and incubation feeding rates should be highest in species using nest sites that are exposed to extreme temperatures that can threaten embryo development or survival. An alternative hypothesis is that incubation feeding is constrained by nest predation; increased trips to the nest are hypothesized to attract the attention of predators. Females can consume more food in one foraging trip off the nest than a male can bring back to the nest in one trip, so the number of trips to the nest can be reduced by females obtaining most of their own food. Therefore, under the nest predation hypothesis, species with a higher risk of nest predation should show increased foraging by females and lower rates of incubation feeding by males. : Reading Comprehension (RC)