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Between 1995 and 2005, the North Atlantic averaged 4.1 major hurricanes (category 3 or stronger) yearly. From 1971 through 1994, however, only 1.5 such hurricanes had swept through each year. According to an analysis by K. Halimeda Kilbourne, this spike in major-hurricane activity reflects a return to normal frequency after a lull in the 1970s and 1980s, rather than the development of a new pattern. Two factors thought to strongly influence hurricane development are wind shear an atmospheric phenomenon in which adjacent layers of air move at different speeds or in different directions-and sea-surface temperature. Strong wind shear tends to rip apart tropical storms before they strengthen into hurricanes, while higher sea-surface temperatures can provide more energy to a forming hurricane. Kilbourne and her colleagues noted that observed variations in wind shear and sea-surface temperature correlate not only with variations in hurricane activity but also with corresponding variations in two biological phenomena. The luminescence of coral growth rings under ultraviolet light reveals periods in which more organic matter was washed from land by heavy storm rains, which are less likely in conditions that favor higher wind shear. And populations of the microorganism G. bulloides in the upper layers of the tropical ocean are more abundant when there is persistent upwelling of relatively cool, nutrient-rich waters. This upwelling occurs as a result of strong trade winds, which also correlate with relatively high wind shear. When these microorganisms die, they become deposited in seafloor sediments, which thus record changes in their abundance. The researchers examined these biological phenomena in samples from North Atlantic areas in which hurricanes form. Based on these observations, they concluded that overall, between 1730 and 2005, the North Atlantic experienced an annual average of 3.25 category-3-or-stronger hurricanes. However, at least six lengthy intervals since 1730 showed increased hurricane activity. Other analyses of long-term natural records bolster these results. Coastal geologist Jeffrey Donnelly studied lake and lagoon sediments to compare the timing of hurricanes during the past 5,000 years with that of El Niños-weather phenomena that increase North Atlantic wind shear-and found that periods with strong, frequent El Niños experienced a lower-than-average number of hurricanes. : Reading Comprehension (RC)